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Lacerta
August 26, 2026 at 09:34 AM
Added the Chapter 2 sequence between the two fights, a preparation table with Thunderwave Sword and Thick Cuirass stats, and the Lightning Stone detour that hands over both
Lacerta is the second Nushi Emma hunts, a swamp-dwelling reptile that rules the flooded ruins of the Ancient Civilization Zone in the Kanto Region. It is fought twice inside Chapter 2, first at level 13 and again at level 16, and it is weak to Lightning in both encounters. Clearing the second fight unlocks the Lacerta's End achievement.
Detail | Value |
|---|---|
Chapter | Chapter 2 |
Region | Kanto Region |
Location | Ancient Civilization Zone |
Levels | 13 (first fight), 16 (second fight) |
Weakness | Lightning |
Status pressure | Poison resistance is worth stacking before the fight |
Achievement | Lacerta's End |
Approach | The reservoir at the end of the zone's two-generator gate puzzle |
Between the fights | A lightning stone has to be found for the control room panel before the lair opens |
Both fights sit inside Chapter 2, and the gap between them is an errand. Lacerta waits at the reservoir at the end of the zone's two-generator gate puzzle, and beating it there only drives it off. The golem in the control room then asks for a lightning stone to feed the control panel. Retracing the route southeast of the Raging River camp and killing the elite golem there yields both the Lightning Stone and the Thunderwave Sword, with a short meeting with Mikoto on the way back. Slotting the stone opens the gates to the lair, one more control panel unlocks the arena inside, and the level 16 rematch starts there. The kill ends with both of them collapsing, and a memory plays out before they wake up back on the walking vessel.
Each victory hands Emma part of Lacerta's power. The poison-themed Dokuzeri line that runs through the rest of the game starts here.
Fight | Level | Rewards |
|---|---|---|
First | 13 | Lacertine Memories, Nushi Skill: Dokuzeri Shower |
Second | 16 | Bloom Art: Dokuzeri: Fang, Blade Art: Dokuzeri |
Attack | How It Works | Response |
|---|---|---|
Bite Combo | Lunges forward into a three-hit bite chain. | Parry every bite. This is the cleanest way to build FP for Koo and fill the stagger gauge. |
Tail Swipe | Turns and swings its tail in a wide arc. | The wind-up is obvious. Parry, then counterattack. |
Toxic Globs | Raises its head and spits three purple globs in succession. | Dodge each projectile rather than parrying, then close the gap. |
Trees from Below | Pools water around itself to slow movement, glows red, then erupts trees out of the ground. | Unparryable. Jump when the red reticle appears, then punish the recovery. |
Swimming Lunge | Submerges, swims at Emma, and surfaces into a single bite. | It cannot be damaged while submerged. Hold the parry for the moment it breaches. |
Poison Gas Charge | Vents poison gas, then charges in a straight line. | Dodge sideways or parry to stagger it. Fire Arrows, Fire Bolts, or Koo's Higanbana: Seeds ignite the gas and cancel the charge. |
Lightning is the lever for the whole fight. The Thunderwave Sword is handed out during the same stretch of the main story after the first kill, which makes the rematch noticeably shorter, and Lightning Arrows and Lightning Bolts cover the gaps when the boss is out of reach. Koo carries the rest: lightning-element skills for stagger pressure, a binding skill to interrupt the heavy attacks, and a healing option so Emma is not spending stimpacks between openings.
The fight breaks into two halves. Lacerta is slow early, which is the window to learn its parry timings in melee and save ammunition. It speeds up in the second half and spends more time in the water, so switch to ranged damage whenever it swims off and only re-engage once it surfaces.
Use Entanglement Overdrive after the bite combo or the tail swipe, when the recovery is longest.
Ignite the poison gas with fire damage instead of dodging it, which turns the charge into free damage.
Clear any adds quickly so the arena stays a one-on-one fight.
Keep an antidote on Koo, since poison stacks fast in the flooded sections.
The rematch is where the loadout matters, because everything worth bringing to it drops during the detour that unlocks it.
Tool | Where It Comes From | What It Does |
|---|---|---|
Thunderwave Sword | Dropped by the elite golem southeast of the Raging River campsite, together with the Lightning Stone | 51 melee attack with the Tempest passive: every slash calls lightning down on nearby foes and adds 6 lightning buildup |
Thick Cuirass | Strongbox by the Tunnel Ruins: Interior camp, found after the meeting with Kunai | 29 defence, four Spirit Stone slots and 60 Poison resistance, which is the resistance this fight actually taxes |
Lightning Arrows and Lightning Bolts | Crafted ammunition, covered under Bow and Ranged Combat | Weakness damage for the stretches Lacerta spends swimming out of reach |
Higanbana: Seeds | Flame-element Koo skill unlocked from Higanbana: Bloom | Ignites the vented poison gas and cancels the charge outright rather than merely dodging it |
Sakura: Javelin | Electric-element Koo skill | Skewers the target with a sakura branch, and a pierced target can be jumped to, which closes the gap when the boss backs into the water |
One habit is worth building early. When Lacerta pools mud and water around itself, the extended jump off Emma's hair clears the pool outright, and the time spent in the air is long enough to drink a stimpack before landing. That turns the attack most likely to end a run into the safest healing window the fight offers.
Lacerta sits second in the Nushi order after Rangifer and before Taurus. The full encounter list, including the paired fights and the recurring Malefact bosses, is on Bosses, and the shared mechanics behind Nushi rewards are covered under Nushi.
The Parry System page covers the timings the slow first half is there to teach, Entanglement Gauge explains the meter the bite combo and tail swipe recoveries are meant to cash in, and Achievements lists Lacerta's End with the rest of the Nushi trophies.